4 2 . 頠 - . , . , , (Dean, Male, 1964). , (Bishui, 1950; Mitchell, 1964; Clark, Thomson, 1964; Sutherland ct al., 1954 .). , 頠 頠 415 蠠 , - N1. , 蠠 - 5032 Ǻ ( 503), 4958 Ǻ ( 4), 5034 5107 Ǻ ( S4), , , : VN2V(g), VN2V(t) NV ( . . 10).